Observations
A large tree up to 50(—70) m tall with a straight and cylindrical bole free of branches for 15—25 m and an average diameter of 55 cm, but occasionally up to 140 cm, thick bark which forms plates and is grey-brown underneath, but scaly and more reddish tinged upwards, and heavy horizontal or ascending branches; needles in pairs, slender but rigid, 16—25 cm long, with persistent basal sheaths; cones solitary or in pairs, almost sessile, cylindrical, 5—11 cm long, after opening twice as thick and ovoid, generally falling off soon; apophysis broadly tetragonal with a smooth, almost depressed umbo; seed small with a deciduous wing of c. 2.5 cm long. Merkus pines of the Asian mainland and the Philippines differ slightly from those of Sumatra: the seedlings have a 'grass stage', the needles are slightly longer, the cones are less cylindrical, and the seeds nearly twice as heavy. Pinus merkusii is locally common in northern Sumatra up to 2000 m altitude. It is the southernmost occurring pine of all pines, and the only one whose natural distribution extends into the southern hemishpere. See also the table on wood properties.
Selected Sources
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[145]Chudnoff, M., 1984. Tropical timbers of the world. Agricultural Handbook 607. USDA Forest Service, Washington, D.C. 464 pp.
[149]Cooling, E.N.G., 1968. Pinus merkusii. Fast growing timber trees of the lowland tropics No 4. Commonwealth Forestry Institute, Oxford. 169 pp.
[153]Corner, E.J.H., 1988. Wayside trees of Malaya. 3rd edition. 2 volumes. The Malayan Nature Society, Kuala Lumpur. 774 pp.
[157]Critchfield, W.B. & Little, E.L., 1966. Geographic distribution of the pines of the world. Miscellaneous Publications 991. USDA Forest Service, Washington, D.C. 97 pp.
[159]Dahms, K.-G., 1982. Asiatische, Ozeanische und Australische Exporthölzer [Asiatic, Pacific and Australian export timbers]. DRW-Verlag, Stuttgart. 304 pp.
[163]Dallimore, W. & Jackson, A.B., 1966. A handbook of Coniferae and Ginkgoaceae. 4th edition. Revised by S.G. Harisson. Edward Arnold Ltd., London. xix + 729 pp.
[224]Farjon, A., 1984. Pines: drawings and descriptions of the genus Pinus. E.J. Brill, Leiden. 220 pp.
[225]Farjon, A., 1990. A bibliography of Conifers. Regnum Vegetabile 122. Koeltz Scientific Books, Königstein. 129 pp.
[230]Ferguson, J.H.A., 1953. Growth and yield of Pinus merkusii in Indonesia. Tectona 43: 21–36.
[283]Granhof, J.J., 1978. Early development of Pinus merkusii Jungh. & de Vriese at high elevation in North Thailand. In: Nikles, D.G., Burley, J. & Barnes, R.D. (Editors): Progress and problems of genetic improvement of tropical forest trees. Proceedings of a joint workshop IUFRO working parties S202-08 and S203-01, Brisbane. 2 volumes. Commonwealth Forestry Institute, Oxford. pp. 694–698.
[284]Granhof, J.J., 1978. Review of Pinus merkusii Jungh. & de Vriese contributions. In: Nikles, D.G., Burley, J. & Barnes, R.D. (Editors): Progress and problems of genetic improvement of tropical forest trees. Proceedings of a joint workshop IUFRO working parties S202-08 and S203-01, Brisbane. 2 volumes. Commonwealth Forestry Institute, Oxford. pp. 714–721.
[288]Greathouse, T.E., 1973. Pilot plantations for quick-growing industrial tree species, Malaysia. Tree improvement in Malaysian conifer plantations. FAO Report No FO: SF/MAL 12, Technical Report 8. 45 pp.
[318]Heyne, K., 1927. De nuttige planten van Nederlands-Indië [The useful plants of the Dutch East Indies]. 2nd ed. 3 volumes. Departement van Landbouw, Nijverheid en Handel in Nederlandsch-Indië, 's-Gravenhage. 1953 pp.
[330]Howcroft, N.H.S., 1978. Progress in preliminary tree improvement and seed production programme with Pinus merkusii Jungh. et de Vriese in Papua New Guinea. In: Nikles, D.G., Burley, J. & Barnes, R.D. (Editors): Progress and problems of genetic improvement of tropical forest trees. Proceedings of a joint workshop IUFRO working parties S202-08 and S203-01, Brisbane. 2 volumes. Commonwealth Forestry Institute, Oxford. pp. 699–706.
[359]Keating, W.G. & Bolza, E., 1982. Characteristics, properties and uses of timbers. Vol. 1. South-East Asia, Northern Australia and the Pacific. Inkata Press Proprietary Ltd., Melbourne, Sydney & London. 362 pp.
[396]Lamprecht, H., 1989. Silviculture in the tropics. Deutsche Gesellschaft für Technische Zusammenarbeit (GTZ), Eschborn. 296 pp.
[417]Lee, Y.H., Engku Abdul Rahman & Chu, Y.P., 1979. The strength properties of some Malaysian timbers. Revised edition. Malaysian Forest Service Trade Leaflet No 34. Malaysian Timber Industry Board, Kuala Lumpur. 107 pp.
[462]Martawijaya, A. et al., 1989. Atlas kayu Indonesia [Indonesian wood atlas]. Vol. 2. Departemen Kehutanan, Badan Penelitian dan Pengembangan Kehutanan, Bogor. 167 pp.
[464]Martini, M., 1982. Mycorrhiza for afforestation with Pinus merkusii and how it influences other pine species. Training course on mycorrhiza. Provisional Report No 12, Malaysia and IFS Sweden. pp. 167–174.
[487]Mindawati, N., 1988. The growth rate of four forest tree species in the alang-alang (Imperata cylindrica Beauv.) area of Jampang Tengal, Sukabumi. Buletin Penelitian Hutan 494: 31–39.
[488]Mirov, N.T., 1967. The genus Pinus. Ronald Press Company, New York. 602 pp.
[517]Nguyen-kha, 1965. Les forêts de Pinus khasya et de Pinus merkusii du Centre-Vietnam. Etude de la dynamique des sols en liaison avec celle de la végétation [Pinus khasya and Pinus merkusii forests of Central Vietnam. Study of the soil dynamics in relation to that of the vegetation]. Faculté de Science, Université de Nancy.
[519]Noerhadi, E. & Wirjodarmodjo, H., 1980. Vegetative propagation of Tectona grandis L. and Pinus merkusii Jungh. & de Vriese using tissue culture. Duta Rimba 6(42): 11–15.
[579]Reyes, L.J., 1938. Philippine woods. Technical Bulletin No 7. Commonwealth of the Philippines, Department of Agriculture and Commerce. Bureau of Printing, Manila. 536 pp. + 88 plates.
[660]Suhardi, 1984. Effect of type inocula and phosphate levels on mycorrhizal formation and initial growth of Pinus merkusii Jungh. & de Vriese seedlings in two types of soils. University of the Philippines, Laguna. Master thesis. 87 pp.
[722]van Alphen de Veer, E.J., 1953. Plantations of Pinus merkusii as a means of reafforestation in Indonesia. Tectona 43: 119–130.
[748]van Steenis, C.G.G.J. & de Wilde, W.J.J.O. (Editors), 1950–. Flora Malesiana. Kluwer Academic Publishers, Dordrecht, Boston, London.
[764]Webb, D.B. et al., 1984. A guide to species selection for tropical and sub-tropical plantations. 2nd ed. Tropical Forestry Papers No 15. Unit of Tropical Silviculture, Commonwealth Forestry Institute, University of Oxford, Oxford. 256 pp.