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Record Number

5720

PROSEA Handbook Number

5(3): Timber trees; Lesser-known timbers

Taxon

Licania Aubl.

Protologue

Hist. pl. Guiane 1: 119, t. 45 (1775).

Family

CHRYSOBALANACEAE

Chromosome Numbers

x = unknown; 2n = unknown

Vernacular Names

Red beam (En). Brunei: kayu agong, sampaluan (Malay), piasau-piasau (Kedayan). Indonesia: kayu besi (Bangka). Malaysia: merbatu kechil (Peninsular), sampaluan, tampaluan (Dusun, Sabah). Philippines: amayan (general), dagingan (Samar-Leyte Bisaya), gapus (Tagbanua). Thailand: khra, khru ra ton (peninsular).

Origin and Geographic Distribution

Licania comprises over 200 species, the vast majority of which occurs in the Neotropics. One species is found in West Africa and three in the Malesian region, with a single timber-producing one: L. splendens (Korth.) Prance (synonyms: Angelesia splendens Korth., Coccomelia nitida (Hook. f.) Ridley, Parinarium philippinense Elmer). The latter occurs in peninsular Thailand, Peninsular Malaysia, Sumatra, West Java (rare), Borneo and the Philippines.

Uses

The strong and durable wood of Licania is used for house posts, railway sleepers and salt-water piling. Because of its toughness it is suitable for tool handles; it is also used for charcoal production.
Though edible, the fruit is not often eaten.

Production and International Trade

Licania timber may occasionally be sold as "merbatu"", together with that of the genera Atuna, Maranthes and Parinari. Supplies, however, are limited.

Properties

Licania yields a heavy hardwood with a density of 880-1130 kg/m3 at 15% moisture content. Heartwood reddish or pale brown with a pink tinge, not clearly demarcated from the paler sapwood; grain straight; texture fine and even; wood with some watered-silk figure on radial surface. Growth rings indistinct, sometimes visible with a hand lens by dense tissue and parenchyma bands twice as far apart; vessels variable in size, from moderately small to moderately large, with oblique tendency, predominantly solitary, tyloses present, occasionally with chalk-white deposits; parenchyma abundant, apotracheal in narrow bands, evenly spaced except near growth ring boundaries, conspicuous; rays very fine, only visible with a hand lens; ripple marks absent.
The wood is apt to split in seasoning unless proper precautions are taken. It is very hard, very strong and tough. It is difficult to work due to the presence of silica, but a smooth finish is obtained with sharp tools. The wood is moderately durable to durable when exposed to the weather and is resistant to marine borer attack.
See also the table on microscopic wood anatomy.

Botany

An evergreen, small to medium-sized tree up to 25 m tall; bole generally straight, cylindrical, up to 80 cm in diameter, with small buttresses; bark surface becoming irregularly cracked and flaking in small pieces, grey-brown, inner bark red-brown; crown rounded, dark green, young foliage covered with silvery silky hairs. Leaves arranged spirally, simple, entire, glabrous when mature; petiole canaliculate; stipules caducous. Inflorescence an axillary or terminal panicle of cymules. Flowers actinomorphic, with a campanulate tomentose receptacle; sepals 5, free; petals 5, about as large as the sepals; stamens 7-10, included and inserted unilaterally on the margin of the disk; ovary inserted at or near the base of the receptacle, 1-locular with 2 ovules, style basal. Fruit a small, stipitate, pear-shaped, bright red, fleshy drupe; endocarp hard, thin, splitting longitudinally upon germination.
In peninsular Thailand flowering takes place from March-April, whereas fruits have been observed in March-July; in Peninsular Malaysia flowering is in April-June. Other sources mention a period of 6-7 months for fruits to ripen. Single trees may not flower for 1-2 consecutive years. Fruits are dispersed by the fruit pigeon Ducula aenea.
The three Malesian species of Licania form the subgenus Angelesia (Korth.) Prance & F. White. The family Chrysobalanaceae is sometimes treated as a subfamily of the Rosaceae.

Ecology

L. splendens is a fairly frequent tree of well-drained, primary and secondary lowland forest on hills and ridges, or sometimes in thickets, up to 400(-800) m altitude. It has been found in a wide variety of forest types including mixed dipterocarp forest, kerangas, peat-swamp forest, freshwater swamp forest, on seashores and rocky locations, but more frequently on sandy soils.

Silviculture and Management

L. splendens is known to coppice well.

Genetic Resources and Breeding

The fairly common occurrence of L. splendens and its reasonably wide geographic distribution suggest that the risk of genetic erosion is small.

Prospects

L. splendens is expected to continue to be used for marine constructions, but increased utilization for other purposes is unlikely as the wood is extremely hard and very difficult to cut and saw.

Literature

[151]Browne, F.G., 1955. Forest trees of Sarawak and Brunei and their products. Government Printing Office, Kuching, Sarawak. xviii + 369 pp.
[163]Burkill, I.H., 1966. A dictionary of the economic products of the Malay Peninsula. 2nd edition. Ministry of Agriculture and Co-operatives, Kuala Lumpur. Vol. 1 (A-H) pp. 1-1240. Vol. 2 (I-Z) pp. 1241-2444.
[198]Cockburn, P.F., 1976-1980. Trees of Sabah. 2 volumes. Sabah Forest Records No 10. Forest Department Sabah, Sandakan.
[209]Corner, E.J.H., 1988. Wayside trees of Malaya. 3rd edition. 2 volumes. The Malayan Nature Society, Kuala Lumpur. 774 pp.
[267]Desch, H.E., 1941-1954. Manual of Malayan timbers. Malayan Forest Records No 15. 2 volumes. Malaya Publishing House Ltd., Singapore. 762 pp.
[341]Flora Malesiana (various editors), 1950-. Kluwer Academic Publishers, Dordrecht, Boston, London.
[436]Heyne, K., 1927. De nuttige planten van Nederlands-Indiƫ [The useful plants of the Dutch East Indies]. 2nd edition, 3 volumes. Departement van Landbouw, Nijverheid en Handel in Nederlandsch-Indiƫ. 1953 pp. (3rd edition, 1950. van Hoeve, 's-Gravenhage/Bandung. 1660 pp.).
[464]Ilic, J., 1990. The CSIRO macro key for hardwood identification. CSIRO, Highett. 125 pp.
[770]Medway, Lord, 1972. Phenology of a tropical rain forest in Malaya. Biological Journal of the Linnean Society 4(2): 117-146.
[830]Ng, F.S.P. & Loh, H.S., 1974. Flowering-to-fruiting periods of Malaysian trees. Malaysian Forester 37: 127-132.
[861]Oey Djoen Seng, 1951. De soortelijke gewichten van Indonesische houtsoorten en hun betekenis voor de praktijk [Specific gravity of Indonesian woods and its significance for practical use]. Rapport No 46. Bosbouwproefstation, Bogor. 183 pp.
[903]Prance, G.T., 1987. Notulae de chrysobalanaceis malesianis praecursoriae. Brittonia 39: 364-370.
[905]Prance, G.T. & White, F., 1988. The genera of Chrysobalanaceae: a study in practical and theoretical taxonomy and its relevance to evolutionary biology. Philosophical Transactions of the Royal Society of London, B. Biological Sciences 320: 1-184.
[933]Research Institute of Wood Industry, 1988. Identification, properties and uses of some Southeast Asian woods. Chinese Academy of Forestry, Wan Shou Shan, Beijing & International Tropical Timber Organization, Yokohama. 201 pp.
[940]Ridley, H.N., 1922-1925. The flora of the Malay Peninsula. 5 volumes. Government of the Straits Settlements and Federated Malay States. L. Reeve & Co., London.
[1039]Smitinand, T. & Larsen, K. (Editors), 1970-. Flora of Thailand. The Forest Herbarium, Royal Forest Department, Bangkok.
[1040]Smythies, B.E., 1965. Common Sarawak trees. Borneo Literature Bureau, South China Morning Post, Hong Kong. 153 pp.
[1048]Soepadmo, E., Wong, K.M. & Saw, L.G. (Editors), 1995-. Tree flora of Sabah and Sarawak. Sabah Forestry Department, Forest Research Institute Malaysia and Sarawak Forestry Department, Kepong.
[1221]Whitmore, T.C. & Ng, F.S.P. (Editors), 1972-1989. Tree flora of Malaya. A manual for foresters. 4 volumes. Malayan Forest Records No 26. Longman Malaysia Sdn. Berhad, Kuala Lumpur & Petaling Jaya.

Author(s)

G.T. Prance

Correct Citation of this Article

Prance, G.T., 1998. Licania Aubl.. In: Sosef, M.S.M., Hong, L.T. and Prawirohatmodjo, S. (Editors): Plant Resources of South-East Asia No 5(3): Timber trees; Lesser-known timbers. PROSEA Foundation, Bogor, Indonesia. Database record: prota4u.org/prosea

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