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Record Number

5896

PROSEA Handbook Number

5(3): Timber trees; Lesser-known timbers

Taxon

Mimusops L.

Protologue

Sp. pl. 1: 349 (1753); Gen. pl., ed. 5: 165 (1754).

Family

SAPOTACEAE

Chromosome Numbers

x = 12; M. elengi: 2n = 24

Vernacular Names

Asian bulletwood (En). Bukal, tanjung (En, Fr). Indonesia: tanjung (general, trade name), karikis (North Sulawesi), tanjung laut (Moluccas). Malaysia: bitis (general, trade name), elengi (general), mengkula (Peninsular), nyatoh batu (Peninsular, Sabah). Philippines: betis (general, trade name), bansalagin (general), kabiki (Tagalog). Burma (Myanmar): kaya. Laos: 'sa koun, phi koun. Thailand: kun (peninsular), kaeo (northern), phikun (central). Vietnam: c[aa]y vi[eef]t, s[eef]n c[as]t.

Origin and Geographic Distribution

Mimusops comprises about 40 species, 20 of which occur in Africa, some 15 in Madagascar, 5 in the Mascarenes and Seychelles and 1, M. elengi L. (synonyms: M. parvifolia R. Br., M. elengi var. parvifolia (R. Br.) H.J. Lam), in Asia and the Pacific. The latter is probably native to India, Sri Lanka, Burma (Myanmar), Indo-China, Thailand and the Andaman Islands, but is commonly planted throughout Malesia towards the Solomon Islands, New Caledonia, Vanuatu and northern Australia, and many other tropical countries.

Uses

The heavy, strong and durable wood of M. elengi is suitable for heavy general construction, bridge building, boat and ship building, marine construction, flooring, bearings, doors and framing. It has also been used for poles and piles, foundation sills, railway sleepers, paving blocks, mine timber, furniture and cabinet work, vehicle bodies, turnery, tool handles, wheelwrighting, walking sticks, weaving shuttles, bobbins, toys, sporting goods and musical instruments. A good-quality veneer and plywood can be manufactured from the wood. It also yields a good "hot"" fuelwood.
M. elengi has fragrant flowers and is often planted as an ornamental in gardens and along roads, including coastal sites. The leaves are used medicinally for headache and sore eyes, and are smoked to cure infections of the nose and mouth. A decoction of the bark, sometimes mixed with the flowers, has been used against fever, diarrhoea, inflammation of the gums and gonorrhoea and, mixed with tamarind bark, as a lotion for skin complaints. The young fruits have been employed in a gargle for treating sprue. The pounded seeds are used to cure obstinate constipation. Fresh flowers yield an oil used as perfume and are also strung in garlands or necklaces for decoration, or placed in linen-cupboards. In Java the flowers have been used against diarrhoea. The seeds yield an oil after pressing which has been used for cooking and illumination. The fruits are edible but reported as tasteless and/or astringent. In India the bark has been use for tanning, but the amount of tannin is low.

Production and International Trade

M. elengi wood is regarded as a commercial wood in Indonesia, but no trade figures are available. It is probably traded in mixed consignments of "bitis"" wood, which comprises many other Sapotaceae genera, such as Madhuca, Palaquium, Payena and Pouteria. M. elengi wood is traded locally in the Philippines. In Burma (Myanmar) and Sri Lanka, plantations of M. elengi have been established.

Properties

M. elengi yields a heavy hardwood with a density of 780-1120 kg/m3 at 15% moisture content. Heartwood deep red or dark red-brown, often with darker streaks, distinct, but not sharply demarcated from the paler 5-7 cm wide sapwood; grain straight, wavy or slightly interlocked; texture very fine to fine and even; occasional watered-silk figure on tangential face of sapwood; taste bitter; wood contains saponin and lathers when rubbed with water. Growth rings indistinct, boundary occasionally indicated by zone without parenchyma; vessels very small to medium-sized, in radial multiples of 2-5(-more), sometimes solitary, tyloses abundant and with dark coloured deposits; parenchyma abundant, apotracheal diffuse-in-aggregates to more or less continuous and closely spaced bands, distinct with a hand lens; rays very fine to moderately fine, usually visible only with a hand lens; ripple marks absent.
Shrinkage of the wood is moderate to very high and it takes about 2 months to air dry boards 3 cm thick from 40% to 15% moisture content. It is liable to end-splitting, warping and surface checking if not carefully seasoned. The wood is very hard, very strong and tough. It is generally considered rather difficult to difficult to work, especially in sawing, due to the presence of silica, but is easier to work when still green; it finishes very well using sharp tools. The sawdust is irritating to nose and throat. The wood is very durable, even when exposed to the weather or in contact with the ground. The heartwood is very resistant to impregnation with preservatives. The wood is reportedly resistant to marine borer attack and to dry-wood termites. The sapwood is susceptible to Lyctus. The gross energy value of the heartwood is 21 340 kJ/kg, that of the sapwood 21 090 kJ/kg.
See also the tables on microscopic wood anatomy and wood properties.

Botany

Evergreen, monoecious, small to medium-sized tree up to 30(-40) m tall; bole often short and divided into several large main branches but sometimes branchless for up to 15(-20) m, up to 100 cm in diameter, buttresses absent or up to 2 m high; bark surface becoming deeply fissured and sometimes peeling off in thin scales, grey, brown or dark red to blackish, inner bark fibrous, pink or reddish, with scanty watery or white sticky latex; crown dense, rounded and spreading, glossy dark green. Leaves alternate or distantly spiral, simple, with wavy, upcurled margins; stipules minute and caducous. Flowers bisexual or functionally male or functionally female, solitary or fasciculate in leaf axils; sepals in 2 whorls of 4; corolla white, fragrant, with a short tube and 8 lobes each divided in three; stamens 8, alternating with 8 staminodes and inserted on the corolla tube; ovary superior, (6-)8-locular with 1 ovule in each cell, style 1. Fruit a 1-2-seeded, ovoid to ellipsoid berry, ripening orange then red. Seedling with epigeal germination; cotyledons emergent, leafy; hypocotyl elongated; all leaves alternate-spiral; taproot strongly developed.
Seedlings and trees are considered to grow slowly, although in the arboretum of the Forest Research Insitute Malaysia, Kepong a 21-year-old individual was 34.5 m tall with a clear bole of 18.6 m and a diameter at breast height of 53.4 cm, i.e. a mean annual diameter increment of 2.5 cm. In Java trees flower and fruit throughout the year. The scented flowers produce nectar at the base of the ovary, and infrequent visits of insects and bats have been observed. Still, pollination is most likely by wind. The stigma is receptive before the pollen is released, stimulating cross-pollination. The seed is known to be dispersed by bats, but monkeys, squirrels and wild pigs probably also eat the fruits.

Image

Mimusops elengi L. – 1, tree habit; 2, flowering twig; 3, flower; 4, stamen and staminode; 5, fruit.

Ecology

M. elengi is fairly common near the sea, but may also be found in rocky locations and inland forest, up to 600 m altitude. It thrives in areas with perhumid or slightly seasonal rainfall types, is usually found in seasonally dry habitats but can stand waterlogging for up to 2 months. It requires a fertile soil.

Silviculture and Management

M. elengi can be propagated by seed or cuttings. Seed can be stored for about 9 months and needs "after-ripening"" during the first month of storage. There are about 2000 dry seeds/kg. Seeds have 70-90% germination in 17-82 days. It is best sown directly in containers. The seedlings can be planted out when 20-30 cm tall. The rooting success of cuttings 10-15 cm long and with a diameter of 0.5-1 cm is 70-90%. It is a shade-tolerant species which retains a full crown and reproduces satisfactorily under fairly dense shade.

Genetic Resources and Breeding

M. elengi is stored in a germplasm collection in Malaysia, chiefly for its medicinal value. Trees may differ markedly in size depending on the site and origin, which offers potential for initiating selection and possibly breeding activities. In the Philippines M. elengi is classified as a vanishing timber tree.

Prospects

Because of its superior wood quality, it is worthwhile starting silvicultural trials with M. elengi. It has good potential for increased utilization in suitable silvicultural schemes. M. elengi is recommended as an ornamental and shade tree in Malaysia.

Literature

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Author(s)

Noorma Wati Haron

Correct Citation of this Article

Haron, N.W., 1998. Mimusops L.. In: Sosef, M.S.M., Hong, L.T. and Prawirohatmodjo, S. (Editors): Plant Resources of South-East Asia No 5(3): Timber trees; Lesser-known timbers. PROSEA Foundation, Bogor, Indonesia. Database record: prota4u.org/prosea

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