Botany
Deciduous or evergreen, medium-sized or rarely large trees up to 30(-50) m tall; bole straight, usually short, cylindrical, up to 100(-180) cm in diameter, with short buttresses in older trees; bark surface smooth, grey-white mottled. Leaves arranged spirally, simple, angular-lobate, palmately veined; stipules present. Flowers axillary, solitary; calyx tubular, with 5 unequal lobes; petals 5, whitish; stamens united into a briefly 5-lobed staminal column bearing sessile wavy anthers from the middle to the apex; ovary superior, 5-locular with many ovules in each cell, style club-shaped, stigma spiralled. Fruit an oblong, ribbed, 5-valved dehiscent capsule, densely lanate inside.
Growth of O. pyramidale trees can be extremely rapid and is according to Koriba's architectural tree model, characterized by orthotropic axes which branch to produce initially equivalent modules. Subsequently one of these modules becomes dominant, constituting one unit of the sympodial trunk. In Peninsular Malaysia growth is too slow to produce the very low-density wood which is sought after for many of its uses. In trials in Indonesia the mean annual increment was 2.0-6.6 cm in diameter and 1.2-3.3 m in height. In Papua New Guinea trees with a diameter of about 20 cm displayed a mean annual diameter increment of 1.7-4.3 cm. In South America mean annual diameter increment is even up to 10 cm and after 10-12 years, when growth stabilizes, trees are generally 20-25 m tall and about 100 cm in diameter. After 12-15 years growth slows down and the trees deteriorate rapidly. Only under specific circumstances may trees reach 50 m in height. In Java and Peninsular Malaysia O. pyramidale flowers throughout the year and is pollinated by bats. Trees start producing viable seed after (2-)3-4 years and dispersal is by wind.
O. pyramidale is highly variable, which is probably why the genus was formerly thought to comprise at least 11 species.
Literature
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