Botany
Deciduous, medium-sized to large trees up to 50 m tall; bole up to 100(-250) cm in diameter, buttresses small to large (up to 4 m high and spreading up to 2 m) or absent; bark surface smooth to rough, fissured or flaky, pale greyish to reddish-brown, inner bark fibrous, hard, usually pinkish or reddish to deep red-brown, sometimes streaked or mottled, with a strong smell of beans. Leaves alternate or opposite, bipinnate with up to 30(-42) pairs of pinnae; leaflets opposite, sessile; petiole and rachis usually with extrafloral nectaries; stipules small, caducous. Flowers in a long-stalked, pendulous, pyriform to clavate, dense head, sterile flowers at base of inflorescence, male ones in middle portion and bisexual ones at apex, 5-merous; calyx long-tubular or funnel-shaped with imbricate lobes; corolla longer than calyx; stamens 10, connate below, shortly exserted; ovary superior, short-stiped, style exserted. Fruit a leathery or woody, stalked, linear to strap-shaped or oblong pod, usually indehiscent, many-seeded, usually several pods together in a pendent infructescence with swollen receptacle. Seeds in 1 row, ellipsoid, with a pleurogram. Seedling with epigeal germination; cotyledons fleshy, peltate; epicotyl with a scale leaf and subsequently bipinnate leaves.
P. speciosa and P. timoriana show a synchronized annual cycle of flowering, fruiting and leaf fall; the trees are without leaves for 2-3 weeks each year. They start flowering when 10-15 m tall, but vegetatively propagated P. speciosa starts flowering and fruiting a few years after planting. The flowering heads are usually pollinated by bats, but are also visited by insects and birds. They produce a foetid odour and a copious nocturnal supply of nectar. Hornbills, monkeys, squirrels, deer, elephants and wild pigs feed on the fruits and probably disperse the seeds. In Java P. timoriana flowers in April-July and usually many fruits are found in June-August. A 24-year-old P. timoriana tree in the arboretum of the Forest Research Institute Malaysia, Kepong had attained only 13.8 m in height and 9.2 cm in diameter.
Parkia is classified in the tribe Parkieae of the subfamily Mimosoideae, together with Pentaclethra, which is confined to tropical Africa and America.
P. timoriana is often cited as P. javanica (Lamk) Merr., especially in Malaysian literature. Although the latter name is older and would therefore have priority, it has been superseded because the correct identity of the species concerned cannot be recovered. The status of P. intermedia Hassk. is uncertain, but it is probably a hybrid between P. speciosa and P. timoriana; it is found almost exclusively in Java. P. sherfeseei Merr. from the Philippines (Mindanao) is possibly conspecific with P. sumatrana.
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