Botany
An evergreen, medium-sized to large tree up to 40(-60) m tall, all parts smelling of garlic; bole columnar, branchless for up to 25 m, up to 80(-150) cm in diameter, often piped, occasionally with small buttresses; bark surface shallowly fissured and flaking into thin rectangles, grey-brown to dark red-brown, inner bark fibrous, purplish-red, inwards with orange flecks. Leaves arranged spirally, simple, entire, exstipulate; petiole swollen distally. Flowers in an axillary, short raceme, 4-5-merous; calyx cup-shaped, margin wavy to toothed; petals reflexed, white, hairy inside; stamens 8 or 10, inserted in pairs about halfway on the petal; ovary superior, imperfectly 3-4-locular with a single ovule in each cell, stigma minutely lobed. Fruit a thinly fleshy, subglobular, 1-seeded, green drupe; endocarp woody with vertical strands. Seedling with hypogeal germination; cotyledons not emergent; hypocotyl not developed; epicotyl with a few scale leaves followed by normal, spirally arranged leaves.
Sapling leaves are alternate-distichous on the branches but arranged spirally on the orthotropic leader. Growth is slow and trees in a large sample in natural forest in Peninsular Malaysia showed an average annual diameter increment of 0.2-0.3 cm. In plantation trials in Peninsular Malaysia the largest trees of 30-33 years old measured 18-29 cm in diameter and 18-21 m in height. In Peninsular Malaysia flowering is in January-July and fruiting more or less throughout the year; in Borneo fruiting is usually in June-September.
Literature
[3]Abdul Rashid b. Hj. A. Malik, 1983. Malaysian timbers - kulim. Malaysian Forest Service Trade Leaflet No 77. Malaysian Timber Industry Board, Kuala Lumpur. 5 pp.
[50]Anonymous, 1970. Daftar nama pohon-pohonan Kapuas Barito (Kalimantan Selatan) [List of tree species in Kapuas-Barito (South Kalimantan)]. Laporan 121. Bagian Eksplorasi dan Botani, Lembaga Penelitian Hutan, Bogor. 90 pp.
[112]Bianchi, A.T.J., 1941. Verslag omtrent de proefbaanopname van 1940 in het boschcomplex Semangoes (Res. Palembang) [Report on the strip survey of 1940 in the Semangus forest (Palembang regency)]. Tectona 34: 286-328.
[151]Browne, F.G., 1955. Forest trees of Sarawak and Brunei and their products. Government Printing Office, Kuching, Sarawak. xviii + 369 pp.
[162]Burgess, P.F., 1966. Timbers of Sabah. Sabah Forest Records No 6. Forest Department, Sabah, Sandakan. xviii + 501 pp.
[163]Burkill, I.H., 1966. A dictionary of the economic products of the Malay Peninsula. 2nd edition. Ministry of Agriculture and Co-operatives, Kuala Lumpur. Vol. 1 (A-H) pp. 1-1240. Vol. 2 (I-Z) pp. 1241-2444.
[193]Chudnoff, M., 1980. Tropical timbers of the world. Forest Products Laboratory, Forest Service, United States Department of Agriculture, Madison. 831 pp.
[198]Cockburn, P.F., 1976-1980. Trees of Sabah. 2 volumes. Sabah Forest Records No 10. Forest Department Sabah, Sandakan.
[218]Dahms, K.-G., 1982. Asiatische, ozeanische und australische Exporthölzer [Asiatic, Pacific and Australian export timbers]. DRW-Verlag, Stuttgart. 304 pp.
[259]den Berger, L.G., 1926. Mechanical properties of Dutch East Indian timbers. Korte Mededeelingen No 12. Proefstation voor het Boschwezen, Buitenzorg. viii + 63 pp.
[261]den Berger, L.G. & Endert, F.H., 1925. Belangrijke houtsoorten van Nederlandsch-Indië, deel I [Important timbers of the Dutch East Indies, part I]. Mededeelingen No 11. Proefstation voor het Boschwezen, Buitenzorg. 136 pp.
[267]Desch, H.E., 1941-1954. Manual of Malayan timbers. Malayan Forest Records No 15. 2 volumes. Malaya Publishing House Ltd., Singapore. 762 pp.
[341]Flora Malesiana (various editors), 1950-. Kluwer Academic Publishers, Dordrecht, Boston, London.
[354]Foxworthy, F.W., 1927. Commercial timber trees of the Malay Peninsula. Malayan Forest Records No 3. Forest Department, Kuala Lumpur. 185 pp.
[436]Heyne, K., 1927. De nuttige planten van Nederlands-Indië [The useful plants of the Dutch East Indies]. 2nd edition, 3 volumes. Departement van Landbouw, Nijverheid en Handel in Nederlandsch-Indië. 1953 pp. (3rd edition, 1950. van Hoeve, 's-Gravenhage/Bandung. 1660 pp.).
[464]Ilic, J., 1990. The CSIRO macro key for hardwood identification. CSIRO, Highett. 125 pp.
[536]Keating, W.G. & Bolza, E., 1982. Characteristics, properties and uses of timbers. Vol. 1. South-East Asia, northern Australia and the Pacific. Inkata Press Proprietary Ltd., Melbourne, Sydney & London. 362 pp.
[553]Kessler, P.J.A. & Sidiyasa, K., 1994. Trees of the Balikpapan-Samarinda area, East Kalimantan, Indonesia. Tropenbos Series 7. The Tropenbos Foundation, Wageningen. 446 pp.
[571]Kloot, N.H. & Bolza, E., 1961. Properties of timbers imported into Australia. Technological Paper No 12. Division of Forest Products, CSIRO, Melbourne. 79 pp.
[677]Lee, Y.H. & Chu, Y.P., 1965. The strength properties of Malayan timbers. Malayan Forester 28: 307-319.
[678]Lee, Y.H., Engku Abdul Rahman bin Chik & Chu, Y.P., 1979. The strength properties of some Malaysian timbers. Malaysian Forest Service Trade Leaflet No 34 (revised edition). Malaysian Timber Industry Board, Kuala Lumpur. 107 pp.
[740]Malaysian Timber Industry Board, 1984. Peraturan pemeringkatan kayu keras gergaji Malaysia [The Malaysian grading rules for sawn hardwood timber]. Ministry of Primary Industries, Kuala Lumpur. 109 pp.
[741]Malaysian Timber Industry Board, 1986. 100 Malaysian timbers. Kuala Lumpur. x + 226 pp.
[757]Martawijaya, A., Kartasujana, I., Kadir, K. & Prawira, S.A., 1992. Indonesian wood atlas. Vol. 2. Forest Products Research and Development Centre, Bogor. 168 pp.
[829]Ng, F.S.P., 1991-1992. Manual of forest fruits, seeds and seedlings. 2 volumes. Malayan Forest Record No 34. Forest Research Institute Malaysia, Kepong. 997 pp.
[831]Ng, F.S.P. & Mat Asri Ngah Sanah, 1991. Germination and seedling records. Research Pamphlet No 108. Forest Research Institute Malaysia, Kepong. 191 pp.
[832]Ng, F.S.P. & Tang, H.T., 1974. Comparative growth rates of Malaysian trees. Malaysian Forester 37: 2-23.
[861]Oey Djoen Seng, 1951. De soortelijke gewichten van Indonesische houtsoorten en hun betekenis voor de praktijk [Specific gravity of Indonesian woods and its significance for practical use]. Rapport No 46. Bosbouwproefstation, Bogor. 183 pp.
[933]Research Institute of Wood Industry, 1988. Identification, properties and uses of some Southeast Asian woods. Chinese Academy of Forestry, Wan Shou Shan, Beijing & International Tropical Timber Organization, Yokohama. 201 pp.
[1023]Sjape'ie, I., 1954. Gewicht en volume van verschillende vrucht- en zaadsoorten [Weight and volume of various fruits and seeds]. Korte mededeling 20A. Bosbouwproefstation, Bogor. 10 pp.
[1038]Smitinand, T., 1980. Thai plant names. Royal Forest Department, Bangkok. 379 pp.
[1040]Smythies, B.E., 1965. Common Sarawak trees. Borneo Literature Bureau, South China Morning Post, Hong Kong. 153 pp.
[1048]Soepadmo, E., Wong, K.M. & Saw, L.G. (Editors), 1995-. Tree flora of Sabah and Sarawak. Sabah Forestry Department, Forest Research Institute Malaysia and Sarawak Forestry Department, Kepong.
[1052]Stadelman, R.C., 1966. Forests of Southeast Asia. Princeton, Memphis, Tennessee. 245 pp.
[1221]Whitmore, T.C. & Ng, F.S.P. (Editors), 1972-1989. Tree flora of Malaya. A manual for foresters. 4 volumes. Malayan Forest Records No 26. Longman Malaysia Sdn. Berhad, Kuala Lumpur & Petaling Jaya.
[1242]Wong, T.M., 1982. A dictionary of Malaysian timbers. Malayan Forest Records No 30. Forest Research Institute Malaysia, Kepong. 259 pp.