Botany
Evergreen shrubs or small to medium-sized trees up to 35 m tall; bole sometimes poorly shaped, branchless for up to 22 m, up to 60(-100) cm in diameter, sometimes with buttresses up to 6 m high; bark surface smooth to fissured or sometimes scaly, pale brown or yellowish-brown to dark brown or dark grey, inner bark fibrous, white to cream with orange to brown or red-brown mottles, with pleasant odour. Leaves decussate, imparipinnate, rachis with 2 glands near the insertion of the petiolules; leaflets 1-11, opposite, petiolulate, serrate, with 2 glands near the base of the blade; stipules interpetiolar. Flowers in an axillary or terminal raceme or panicle, 5-merous; sepals free; petals free, yellowish-green to greenish-white; stamens with broad filaments; disk annular; ovary superior, (2-)3(-4)-locular with 1-many ovules in each cell, styles closely appressed, forming 1 peltate stigma. Fruit a globose to 3-lobed, indehiscent berry, crowned by radial lines or horn-like structures. Seed mostly angular, with endosperm. Seedling with epigeal germination; cotyledons emergent; hypocotyl elongated.
In Borneo most species flower in November-May; in Java T. sphaerocarpa has been recorded flowering throughout the year; in the Philippines T. ovalifolia flowers from December to February. The disk of the sweet-scented flowers produces honey, probably attracting insects for pollination. Fruits take about 5 months to mature. The fruits are probably eaten and dispersed by birds and other animals.
Literature
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